Acknowledgements
This study was financially supported by the Hainan Province Science and Technology Special Fund (No. ZDYF2021XDNY142 YSPTZX202018); the National Natural Science Foundation of China (NSFC, No. 31660033); the Hainan Provincial Natural Science Foundation of China (No. 321RC470); and the Hainan Yazhou Bay Seed Laboratory (No. B21HJ0905). We thank LetPub (www.letpub.com) for linguistic assistance and pre-submission expert review.