AUTHOR CONTRIBUTIONS
J.P.C., A.J.W., D.P.L.T., T.J.B., and J.P.P. conceived the study and performed field work. D.P.L.T. and A.W.W. processed genetic samples and conducted statistical analyses; J.P.C., A.J.W., D.P.L.T., T.J.B., A.W.W., and J.P.P. wrote and reviewed the manuscript.