Silverthorn, T.K. * and Richardson, J.S. 
Department of Forest and Conservation Sciences, The University of British Columbia, 2424 Main Mall, Vancouver, BC V6T 1Z4, Canada
* Corresponding author
Email: teresa.silverthorn@alumni.ubc.ca; teresa.silverthorn@gmail.com 
A version of the manuscript is published at: https://doi.org/10.1007/s10021-021-00621-z