Extraction with Traitor software
The Traitor software https://github.com/TankredO/traitor was used to segment, align, and extract morphological traits from images (Dayrell et al. in press). The extracted traits include morphometric measurements (length, width, aspect ratio, area, perimeter, diaspore surface structure, solidity, circularity”), colour measurements (sRGB, linear sRGB, dominant colours), and 50 coordinates of the shape outline standardised for size invariance.