The Traitor software
https://github.com/TankredO/traitor
was used to segment, align, and extract morphological traits from images
(Dayrell et al. in press). The extracted traits include
morphometric measurements (length, width, aspect ratio, area, perimeter,
diaspore surface structure, solidity, circularity”), colour measurements
(sRGB, linear sRGB, dominant colours), and 50 coordinates of the shape
outline standardised for size invariance.