\documentclass[10pt]{article}

\usepackage{fullpage}
\usepackage{setspace}
\usepackage{parskip}
\usepackage{titlesec}
\usepackage[section]{placeins}
\usepackage{xcolor}
\usepackage{breakcites}
\usepackage{lineno}
\usepackage{hyphenat}





\PassOptionsToPackage{hyphens}{url}
\usepackage[colorlinks = true,
            linkcolor = blue,
            urlcolor  = blue,
            citecolor = blue,
            anchorcolor = blue]{hyperref}
\usepackage{etoolbox}
\makeatletter
\patchcmd\@combinedblfloats{\box\@outputbox}{\unvbox\@outputbox}{}{%
  \errmessage{\noexpand\@combinedblfloats could not be patched}%
}%
\makeatother


\usepackage{natbib}




\renewenvironment{abstract}
  {{\bfseries\noindent{\abstractname}\par\nobreak}\footnotesize}
  {\bigskip}

\titlespacing{\section}{0pt}{*3}{*1}
\titlespacing{\subsection}{0pt}{*2}{*0.5}
\titlespacing{\subsubsection}{0pt}{*1.5}{0pt}


\usepackage{authblk}


\usepackage{graphicx}
\usepackage[space]{grffile}
\usepackage{latexsym}
\usepackage{textcomp}
\usepackage{longtable}
\usepackage{tabulary}
\usepackage{booktabs,array,multirow}
\usepackage{amsfonts,amsmath,amssymb}
\providecommand\citet{\cite}
\providecommand\citep{\cite}
\providecommand\citealt{\cite}
% You can conditionalize code for latexml or normal latex using this.
\newif\iflatexml\latexmlfalse
\providecommand{\tightlist}{\setlength{\itemsep}{0pt}\setlength{\parskip}{0pt}}%

\AtBeginDocument{\DeclareGraphicsExtensions{.pdf,.PDF,.eps,.EPS,.png,.PNG,.tif,.TIF,.jpg,.JPG,.jpeg,.JPEG}}

\usepackage[utf8]{inputenc}
\usepackage[english]{babel}



\usepackage{float}








\begin{document}

\title{Effects of population density and environmental conditions on
life-history prevalence in a migratory fish}



\author[1]{Mark Sorel}%
\author[2]{Andrew Murdoch}%
\author[3]{Richard Zabel}%
\author[4]{Cory Kamphaus}%
\author[5]{Eric Buhle}%
\author[6]{Mark Scheuerell}%
\author[7]{Sarah Converse}%
\affil[1]{University of Washington}%
\affil[2]{Washington Department of Fish and Wildlife}%
\affil[3]{National Marine Fisheries Service}%
\affil[4]{Yakama Nation Fisheries}%
\affil[5]{NOAA Fisheries Northwest Fisheries Science Center}%
\affil[6]{NOAA Fisheries Service}%
\affil[7]{University of Washington School of Aquatic and Fishery Sciences}%


\vspace{-1em}



  
  \date{May 26, 2024}


\begingroup
\let\center\flushleft
\let\endcenter\endflushleft
\maketitle
\endgroup





\selectlanguage{english}
\begin{abstract}
1) Individual variation in life-history traits can have important
implications for the ability of populations to respond to environmental
variability and change. 2) In migratory animals, flexibility in the
timing of life-history events, such as juvenile emigration from natal
areas, can influence the effects of population density and environmental
conditions on habitat use and population dynamics. 3) We evaluated the
functional relationships between population density and environmental
covariates and the abundance of juveniles expressing different
life-history pathways in a migratory fish, Chinook salmon (Oncorhynchus
tshawytscha), in the Wenatchee River basin in Washington State, USA. 4)
We found that the abundance of younger emigrants from natal streams was
best described by an accelerating or near-linear function of spawners,
whereas the abundance of older emigrants was best described by a
decelerating function of spawners. This supports the hypothesis that
emigration timing varies in response to density in natal areas, with
younger-emigrating life-history pathways comprising a larger proportion
of emigrants when densities of conspecifics are high. 5) We also
observed positive relationships between winter stream discharge and
abundance of younger emigrants, supporting the hypothesis that habitat
conditions can also influence the prevalence of different life-history
pathways. 6) Our results suggest that early emigration, and a resultant
increase in the use of downstream rearing habitats, may increase at
higher population densities and with greater winter precipitation.
Winter precipitation is projected to increase in this system due to
climate warming. 7) Characterizing relationships between life-history
prevalence and environmental conditions may improve our understanding of
species habitat requirements and is a necessary first step in
understanding the dynamics of species with diverse life-history
strategies. 8) As environmental conditions change -- due to climate
change, management, or other factors -- resultant life-history changes
are likely to have important demographic implications that will be
challenging to predict if life-history diversity is not accounted for in
population models.%
\end{abstract}%



\sloppy




\selectlanguage{english}
\FloatBarrier
\end{document}

