Author Contributions

Fortunate M. Phaka carried out fieldwork, conducted laboratory analysis and wrote the initial draft of the manuscript; Fortunate M. Phaka, Louis H. du Preez, Maarten P. M. Vanhove, Jean Hugé: designed the project, acquired funding, and revised the initial manuscript; Edward Netherlands, Maarten Van Steenberge provided training and supervision for laboratory analysis, revised the initial manuscript; Erik Verheyen , Gontran Sonet contributed to the project design, and revised the initial manuscript.