Author Contributions
Fortunate M. Phaka carried out fieldwork, conducted laboratory analysis
and wrote the initial draft of the manuscript; Fortunate M. Phaka, Louis
H. du Preez, Maarten P. M. Vanhove, Jean Hugé: designed the project,
acquired funding, and revised the initial manuscript; Edward
Netherlands, Maarten Van Steenberge provided training and supervision
for laboratory analysis, revised the initial manuscript; Erik Verheyen ,
Gontran Sonet contributed to the project design, and revised the initial
manuscript.