Habitat fragmentation and landscape modification are also severe threats to terrestrial biodiversity (Fischer & Lindenmayer 2007) and the linkages between terrestrial and freshwater ecosystems. For example, artificial constructions such as roads or fences have been shown to obstruct long-distance dispersal in mammals (Seidler et al. 2015; Tucker et al. 2018; Bartoń et al. 2019), plants (Deneret al. 2021) and even microbes (Le Provost et al. 2021), which were historically considered to be near-universally distributed. Interestingly, some freshwater species also utilize the terrestrial matrix for dispersal, which can help these taxa overcome effects of habitat fragmentation (Zuluaga et al. 2022). For example, freshwater invertebrates with adults dispersing actively over land through the air better track environmental variation and are less affected by barriers such as dams (Grönroos et al. 2013; Tonkinet al. 2018). Similarly, the ability of freshwater organisms to use other organisms (e.g., birds) as dispersal vectors may both mitigate effects of habitat fragmentation and facilitate the spread of invasive species (Incagnone et al. 2015; Coughlan et al. 2017).