Acknowledgements

We thank Dr. Francesco Lamanna and Dr. Christiane Paul for participation in sample collection in Congo/Brazzaville and Dr. Victor Mamonekene, University of Brazzaville, for logistical support during field work. Support from Katrin Kiemel in data analysis and manuscript revision is also acknowledged. Financial support for this study was provided by the University of Potsdam. R. Amen was funded by the Bridge Scholarship Program of the University of Potsdam and by the Ministry of Higher Education of the Arab Republic of Egypt.