Table 2 – Estimated parameters (β-est), standard errors (SE), t-values (t) and p-values (P) for each covariate in the GLMMs performed on: a) ASV-ratios, b) β- replacement, and c) β- richness of the three examined groups. Dist_path = ordinal variable representing increasing distance from the tourist path; Dist_entrance = ordinal variable representing increasing distance from the tourist path; substrate = dissimilarity of the substrate composition from the control area. Significant results are highlighted in bold.