ACKNOWLEDGMENTS
This work was financially supported by the National Key Research Development Program of China (No.2022YFC2105403), the National Natural Science Foundation of China (No. 21276081), the Scientific Research Think Tank of Biological Manufacturing Industry in Qingdao (QDSWZK202004), the Fundamental Research Funds for the Central Universities (JKF01211116, JKVF1211036, JKF01211708).