Figure 4. Trait covariance along environmental gradients.Environment across sites was characterized using Climatic Water Deficit, soil C:N ratio, and two composite climate axes derived from Principal Components Analysis of selected climate variables from WorldClim data (see Methods). Only relationships with significant trends are shown here. See Table S1 for all coefficient estimates. Prior to analysis, all predictors were standardized by subracting from their mean and dividing by one standard deviation and back transformed to the original scale.