ACKNOWLEDGEMENTS
The study was supported by the National Natural Science Foundation of China (U20A2011; 41930652; 32001215; 31870502; 31870503); CAS Key Laboratory of Agro-ecological Processes in Subtropical Region, Institute of Subtropical Agriculture (ISA2020104); the China Postdoctoral Science Foundation (2020M682586); and Guangxi Natural Science Foundation Program (2020JJB130003; 2020GXNSFDA238012).