Acknowledgements
We are grateful to members of the
Department of Population biology for comments on the previous version of
the manuscript. This research was supported by the Czech Science
Foundation project GAČR 17-11281S and partly by the institutional
research project RVO 67985939. JXQ
was financially supported by the National Natural Science Foundation of
China (31200249), National Special Program on Basic Works for Science
and Technology of China (2015FY1103003-6).