Acknowledgments
This study was financially supported by the National Research Foundation of Korea (2017R1A2B3004972, NRF-2020R1F1052024), the Research Institute of Veterinary Science and the BK21 PLUS Program for Creative Veterinary Science Research, and a SNU-Grant (#550–2020005). We also thank members of the G. Jang lab for their valuable comments and the technical support of SR Han and SY Yum