Author contributions
JS, PT, PL, and AF conceived the study. JS, AF, PT, DB, CS and PL
participated in the sampling of fishes and collection of DNA samples.
JS, YY and DB performed the laboratory work. JS and YY performed the
bioinformatical analyses. All authors contributed to interpretation of
results. JS, YY and AF drafted the first manuscript. All authors
commented on and agreed to the final version of the manuscript.