Acknowledgements
We are grateful the reviewers and editors who provided useful feedback on this manuscript. Funding for this research was provided by the National Natural Science Foundation of China (NSFC, 31800335, 32171511), Guizhou Province Science and Technology Department (2019-1089), Ecology Domestic First-class Discipline Construction Program (GNYL2017-007) and Guizhou University Talent Introduction Research Program (2017-39) to SW.