ACKNOWLEDGEMENTS
We thank the University of Ibagué and University of Tolima for supporting this research. We are grateful to the Departamento Administrativo de Ciencia, Tecnología e Innovación (Colciencias-Currently Minciencias, Colombia) for financial support for this project (ID Project 130780864623). The first author thanks to Oficina de Investigaciones y Desarrollo Científico de la Universidad del Tolima by the early researcher scholarship. We thanks to Roberto Builes, Alexander Cuervo, Eliana Loaiza and Francisco Chávarro by their field work assistance. Specimens collected are covered through the Colombian “Permiso Marco de Recolección”, granted to the Universidad de Ibagué (Resolución no. 01003, 07 June 2019, Autoridad Nacional de Licencias Ambientales –ANLA). All laboratory procedures were followed in agree the Ethical Code approved for the above Minciencias’ project. AvdM was financed through FCT – Fundação para a Ciência e a Tecnologia, I.P. under contract number DL57/2016/CP1440/CT0009.