Acknowledgements
ACH was supported by Chinese National Natural Science Foundation (Grant #:U1602265, Mapping Karst Biodiversity in Yunnan), the Strategic Priority Research Program of the Chinese Academy of Sciences (Grant No.XDA20050202), the High-End Foreign Experts Program of Yunnan Province (Grant #:Y9YN021B01, Yunnan Bioacoustic monitoring program), the CAS 135 program (No. 2017XTBG-T03), and the Chinese Academy of Sciences Southeast Asia Biodiversity Research Center fund (Grant #:Y4ZK111B01). MCO was supported by the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB310304), the NSFC International Young Scholars Program (31850410464) and the CAS President’s International Fellowship Initiative (2018PB0003, 2020PB0142). HJQ was supported by the NSFC (31772432).