ACKNOWLEDGEMENTS
We thank Albina Ilyasova, Wenli Ding, Yunhe Wang, Asad Prodhan and Patrick E. Hayes for their assistance in aspects of this work, and Rob Creasy and Bill Piasini for help with maintaining the plants in the glasshouse. Z.H. was supported by China Scholarship Council Project grant (201508220115). This research was supported by an Australian Research Council Discovery Project grant (DP130100005) awarded to H.L.