Acknowledgments
This work was supported by grants from the US National Science Foundation Long-Term Ecological Research Program (LTER) including DEB-1234162 and DEB-1831944. Funding from the UMN Graduate School, from EEB and Cedar Creek supported MK. We are grateful to Troy Mielke, Jon Anderson, Anita Krause, Susan Barott and Dan Bahauddin for logistical support, Sarah Hobbie and Forest Isbell for comments on the manuscript, Evelyn Hayhurst, Katherine Schroeder, Megan Wilcots, Lucas Bowerman and Cedar Creek interns for their help in data collection.