Acknowledgements
We are grateful to all contributors who assisted with sampling
endeavors, especially Mauricio Jimenez, Maria Isabel DiMare, Sofia
Bernal, Genuar Nuñez, Natalia Valverde and Mauricio Losilla. We would
also like to thank the National System of Conservation Areas (SINAC),
and the Ministry of Environment and Energy (MINAE) of Costa Rica. This
research was carried out thanks to funding by the Cleveland Zoological
Society and Cleveland Metroparks Zoo.
Ethics statement: The present study was conducted under the
protocols established by the Institutional Committee for the Care and
Use of Animals (Comité Institucional para el Cuidado y Uso de los
Animales) of the Universidad de Costa Rica, adhered to the legal
requirements of Costa Rica, and adhered to the American Society of
Primatologists (ASP) Principles for the Ethical Treatment of Non-Human
Primates. Collection permit number: MINAET-SINAC-Costa Rica:
042-2012-SINAC.
Data availability statement: The data that support the findings
of this study are available from the corresponding author upon
reasonable request.
Author contributions statement: C.N., and A.C. conceived the
investigation and wrote the article, A.C., and G.G-E collected samples,
C.N., N.R., and C.S., conducted the laboratory analysis, C.N., M. S.,
C.I-C., A.C. and O.R-C.: performed the statistical analysis, J.F., and
G.S. copy reviewed and edited. All authors reviewed the manuscript.
Conflict of interest statement: The author(s) declare no
competing interests.