Acknowledgements
We are grateful to all contributors who assisted with sampling endeavors, especially Mauricio Jimenez, Maria Isabel DiMare, Sofia Bernal, Genuar Nuñez, Natalia Valverde and Mauricio Losilla. We would also like to thank the National System of Conservation Areas (SINAC), and the Ministry of Environment and Energy (MINAE) of Costa Rica. This research was carried out thanks to funding by the Cleveland Zoological Society and Cleveland Metroparks Zoo.
Ethics statement: The present study was conducted under the protocols established by the Institutional Committee for the Care and Use of Animals (Comité Institucional para el Cuidado y Uso de los Animales) of the Universidad de Costa Rica, adhered to the legal requirements of Costa Rica, and adhered to the American Society of Primatologists (ASP) Principles for the Ethical Treatment of Non-Human Primates. Collection permit number: MINAET-SINAC-Costa Rica: 042-2012-SINAC.
Data availability statement: The data that support the findings of this study are available from the corresponding author upon reasonable request.
Author contributions statement: C.N., and A.C. conceived the investigation and wrote the article, A.C., and G.G-E collected samples, C.N., N.R., and C.S., conducted the laboratory analysis, C.N., M. S., C.I-C., A.C. and O.R-C.: performed the statistical analysis, J.F., and G.S. copy reviewed and edited. All authors reviewed the manuscript.
Conflict of interest statement: The author(s) declare no competing interests.