Acknowledgements
This research was supported by grants from the National Research Foundation (NRF) of South Africa (Grants 91599 and 98836), the Palaeontological Scientific Trust (PAST), and the Centre for Excellence in Palaeosciences, University of the Witwatersrand. Fieldwork procedures were approved by the Animal Ethics Committee of the University of the Free State (experiment number 08/2014), and permission granted from the Department of Economic Development, Tourism and Environmental Affairs, Free State Province (permit 01/25892), who also provided population census data for the eight provincial protected areas (with thanks to Erika Schulze). We acknowledge assistance from Izak Thapo, Adam Thibeletsa†, Jacob Maine, Bonny Nduma†, and Abel Dichakane in the field, Grant Hall with stable isotope analysis, and especially James Brink† for sharing his unparalleled insights into the natural history of African mammals in discussions over many years.