Acknowledgements
This research was supported by grants from the National Research
Foundation (NRF) of South Africa (Grants 91599 and 98836), the
Palaeontological Scientific Trust (PAST), and the Centre for Excellence
in Palaeosciences, University of the Witwatersrand. Fieldwork procedures
were approved by the Animal Ethics Committee of the University of the
Free State (experiment number 08/2014), and permission granted from the
Department of Economic Development, Tourism and Environmental Affairs,
Free State Province (permit 01/25892), who also provided population
census data for the eight provincial protected areas (with thanks to
Erika Schulze). We acknowledge assistance from Izak Thapo, Adam
Thibeletsa†, Jacob Maine, Bonny Nduma†, and Abel Dichakane in the field,
Grant Hall with stable isotope analysis, and especially James Brink† for
sharing his unparalleled insights into the natural history of African
mammals in discussions over many years.