Acknowledges

We thank Wenjun Teng, Chao Han, and Ruibin Xue for help with the field work. This study was supported by Beijing Natural Science Foundation (Grant No. 5204030), National Natural Science Foundation of China (Grant No. 31901173), and Beijing Academy of Agriculture and Forestry Sciences (BAAFS) Special Project on Hi-Tech Innovation Capacity (Grant No. KJCX20180707 and KJCX20200527).