Acknowledgements
We thank the researchers whose published data we used in this paper. Thanks to Robert Ricklefs, Christine Edwards, and Carmen Ulloa for advice in this study. Isabel Loza, Justin Baldwin and Sebastián Tello provided valuable help with statistical analyses. Many thanks to Justin Zweck, Justin Baldwin, Krissa Skogen, and to members of the Muchhala lab at the University of Missouri at Saint Louis for constructive discussions on a previous version of this manuscript. This research was supported by the Whitney Harris World Ecology Center at the University of Missouri at Saint Louis.